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Japanese macaques form matrilineal troops that usually range from 20 to 30 individuals, but can sometimes be as large as over 100. A major constraint on troop size is food availability. The troop is ruled by an alpha female and an alpha male. Females typically outnumber males in the troop 3 to 1 and are ranked by a hierarchy that is inherited and passed from mother to infant. The alpha male is responsible for fathering the offspring of the group as well as providing protection and leading the movement of the group.